Prehistory

notes
vsi
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Human history before the written record including timelines and clades of human evolution.
Author

Stephen J. Mildenhall

Published

2025-04-08

Modified

2025-04-08

Introduction

Prehistory covers the period before the start of written history and includes the emergence of Homo sapiens as a species. Notes motivated by the Prehistory VSI.

Timelines

Stone Age, Bronze Age, and Iron Age

The Three Age system had its uses in the 19th and 20th centuries [but] divisions of time masked continuities between periods.

Comments about the over-simplistic nature of the Three Age system notwithstanding, Table 1 gives a timeline from the dawn of humanity to the rise of the Egyptian Empire, using archaeological periods common in Europe and the Near East. Dates vary by region. These are approximate and Near East-centric when relevant. Interspersed are dates of the Last Glacial Maximum and the rise of the Egyptian empire. Notice how the dates compress to more recent times. Statistically, it’s all Lower Paleolithic!

Table 1: Timeline from the dawn of humanity to the rise of the Egyptian Empire.
Period Approximate Years Ago Key Developments
Lower Paleolithic >3,000,000 – 300,000 ya First stone tools (Oldowan, then Acheulean); Homo habilis, Homo erectus.
Middle Paleolithic 300,000 – 47,000 ya Neanderthals in Europe, early Homo sapiens in Africa; Mousterian tools.
Upper Paleolithic 47,000 – 12,000 ya Homo sapiens spreads globally; art (e.g., cave paintings), advanced tools.
Last Glacial Maximum (LGM) ~26,500 to ~19,000 ya Coldest point in the last ice ace
De-glaciation ~19,000 to ~11,700 ya
Mesolithic 12,000 – 10,000 ya Transition to farming; microlithic tools; regional.
Start of Holocene epoch 11,700 ya 9,750 BCE, end of last major glaciation, rapid warming
Neolithic 10,000 – 6,000 ya Farming, permanent villages, pottery, early religion.
PPNB mega-sites from 10,000 ya See Table 2 for details
Chalcolithic (Copper Age) 7,000 – 5,300 ya Use of copper tools; proto-urban settlements (e.g., Çatalhöyük).
Bronze Age 5,300 – 3,200 ya Metal alloys (bronze); writing (cuneiform, hieroglyphs); early states.
Stonehenge started 5,000 ya
Old Kingdom Egypt ~4,700 ya begins around 2,686 BCE, Egyptian Empire rises
Great Pyramid of Giza 4,500 ya
Iron Age 3,200 – 1,400 ya Iron tools; rise of empires (Assyria, Babylon, Persia).

The Holocene is the current geological epoch, beginning 11,700 years ago at the end of the last major glaciation. It is formally defined by a sharp warming event recorded in the Greenland NGRIP ice core, where a marked rise in deuterium levels signals the end of the Younger Dryas, a brief return to colder conditions following the Last Glacial Maximum. This transition marks the start of the present interglacial period, characterized by relatively stable and warm climates in which agriculture and human civilization developed.

PPNB Mega-Sites and Other Significant Constructions

Table 2 is a list of major PPNB (Pre-Pottery Neolithic B) mega-sites, with a few key facts for each. These sites are mostly from the Levant and Anatolia (most of modern-day Turkey), dated roughly 10,700–8,200 ya (8,700–6,200 BCE).

Table 2: Major PPNB mega-sites.
Site Region Dates (ya) Key Facts Structures Est. Population
Aşıklı Höyük Central Anatolia 10,400–9,300 Early domestic sheep/goats; burned dwellings; planned layout ~400 buildings 400–1,000
Çayönü Southeastern Turkey 10,300–9,000 Copper use; architectural diversity; skull cults >200 structures 500–1,000
Göbekli Tepe Southeastern Turkey 11,500–10,000 T-shaped pillars; ritual focus; no dwellings ~20 circular enclosures None permanent (ritual site)
Jericho (Tell es-Sultan) West Bank 11,000–9,000 Oldest city; stone tower and wall Hundreds over time Up to 2,000
Ain Ghazal Jordan 9,600–8,200 Plaster statues; signs of overuse of environment ~150 structures 2,000–3,000
Beidha Southern Jordan 10,200–9,200 Planned village; early herding; round to rectangular buildings ~65 buildings 200–300
Abu Hureyra Syria 13,000–7,000 Domestication sequence; dietary shifts Hundreds across phases Several hundred to 2,000
Nevalı Çori Southeastern Turkey 10,500–9,000 Pre-Göbekli architecture; skull caching ~30 buildings 100–200

Stonehenge was constructed in phases between about 5,000 and 4,000 years ago, with the earliest earthworks dating to around 5,100 years ago and the main stone circle erected around 4,500 years ago. It is part of a broader ritual landscape that includes Durrington Walls, a large Neolithic settlement, and Avebury, the largest stone circle in Europe, built around 4,600 years ago. Other major sites include Silbury Hill, a massive artificial mound from about 4,400 years ago; Grime’s Graves, a Neolithic flint mine complex in Norfolk; and the Ring of Brodgar and Maeshowe on Orkney, part of a dense ceremonial complex dating to about 5,000–4,500 years ago. These are considered true prehistoric mega-sites in the British Isles.

The Great Pyramid of Giza was built around 4,500 years ago, during the reign of Pharaoh Khufu of Egypt’s Fourth Dynasty. It is the largest of the three pyramids on the Giza Plateau and originally stood about 146.6 meters tall. Constructed as a royal tomb, it was part of a larger complex that included subsidiary pyramids, temples, and causeways. The two other major pyramids at Giza, built for Khafre and Menkaure, followed within the next century. These pyramids are the best-known and most enduring monuments of Egypt’s Old Kingdom, a period that lasted from about 4,700 to 4,200 years ago, when pyramid-building reached its height.

These sites are shown in the nearby map. The map was created with the code in Section 6.

Hominins

Hominins are all species on the human side of the human–chimpanzee split.

A hominin is any species that belongs to the group consisting of modern humans (Homo sapiens), our immediate ancestors, and all extinct relatives more closely related to us than to chimpanzees. This includes species in the genera Homo, Australopithecus, Paranthropus, and Ardipithecus.

Monkeys and Apes

Monkeys are primates that typically have tails, walk on all fours, and are generally smaller and more arboreal than apes. They include New World species like capuchins and howlers, and Old World species like baboons and macaques. Apes, by contrast, are tailless, usually larger, and have a more upright posture, with advanced cognitive and social abilities. The apes include gibbons, orangutans, gorillas, chimpanzees, and humans. While both groups are part of the primate order, apes belong to the Hominoidea clade and are more closely related to humans than monkeys.

Humans in Context within Primates

The next diagram is a cladogram for humans, chimps, gorillas, orangutans, baboons, and lemurs, including approximate divergence dates. It is a text-based cladogram with nested clades (monophyletic groups).

Primates (~85–90 million years ago)
├── Strepsirrhini (~63–85 million years ago)
│   └── Lemuriformes (~60 Mya)
│       └── Lemuridae (~50 Mya)
│           └── Lemur, Eulemur, etc.              ← lemurs
└── Haplorhini (~63–70 Mya)
    ├── Platyrrhini (New World monkeys, ~40–45 Mya)
    │   ├── Cebus                                 ← capuchins
    │   ├── Alouatta                              ← howlers
    │   └── Saguinus, Callithrix, etc.            ← tamarins, marmosets
    └── Catarrhini (Old World monkeys + apes, ~25–30 Mya)
        ├── Cercopithecoidea (Old World monkeys, ~25 Mya)
        │   ├── Papio                              ← baboons
        │   ├── Macaca                             ← macaques
        │   └── Colobus, Cercopithecus, etc.       ← colobines, guenons
        └── Hominoidea (apes, ~25 Mya)
            ├── Hylobatidae (~18–20 Mya)
            │   └── Hylobates, Nomascus, etc.     ← gibbons
            ├── Ponginae (~16–18 Mya)
            │   ├── Sivapithecus †                 ← Miocene relative of orangutans
            │   └── Pongo                          ← orangutans
            └── Homininae (~8–10 Mya)
                ├── Gorilla                        ← gorillas
                └── Hominini (~6–7 Mya)
                    ├── Pan                        ← chimpanzees, bonobos
                    └── Homo lineage
                        ├── Sahelanthropus †       ← ~7 Mya
                        ├── Ardipithecus †         ← ~5–6 Mya
                        ├── Australopithecus †     ← ~4–2 Mya
                        ├── Paranthropus †         ← ~2.7–1.2 Mya
                        └── Homo
                            ├── Homo habilis †      ← ~2.4–1.6 Mya
                            ├── Homo erectus †      ← ~2–0.1 Mya
                            ├── Homo neanderthalensis † ← ~400–40 kya
                            ├── Homo floresiensis † ← ~100–50 kya
                            └── Homo sapiens        ← modern humans, ~300 kya

Figure 1 and Figure 2 show more detailed cladograms from two recent papers in the field.

Figure 1: Primate clade. Source: [1].
Figure 2: Primate clade and images. Source: [2],

Major Clades

Clade Description Members from your list
Primates All listed species All
Strepsirrhini “Wet-nosed” primates Lemurs
Haplorhini “Dry-nosed” primates All others
Anthropoidea (Simiiformes) Monkeys & apes Humans, chimps, gorillas, orangutans, baboons, gibbons
Catarrhini Old World monkeys & apes Humans, chimps, gorillas, orangutans, baboons, gibbons
Hominoidea Apes and humans Humans, chimps, gorillas, orangutans, gibbons
Hylobatidae Lesser apes Gibbons
Hominidae Great apes & humans Humans, chimps, gorillas, orangutans
Homininae African great apes Humans, chimps, gorillas
Hominini Humans and closest relatives Humans, chimps
Homo Genus Homo Humans, extinct hominins

Note: Anthropoidea (or Simiiformes) is older terminology and less commonly used in modern cladistics. It includes all monkeys and apes, Platyrrhini (New World monkeys) and Catarrhini (Old World monkeys + apes). It excludes Strepsirrhini (lemurs, lorises) and Tarsiers, so it’s a proper clade under Haplorhini. In cladistic usage today, people more often say Haplorhini → Platyrrhini + Catarrhini, and skip over “Anthropoidea” entirely.

Approximate Divergence Timeline

Split Clades ~Date (Mya)
Lemurs from rest Strepsirrhini vs Haplorhini ~63–85 Mya
New World monkeys from others Platyrrhini vs Catarrhini ~40 Mya
Old World monkeys vs apes Cercopithecoidea vs Hominoidea ~25 Mya
Gibbons split Hylobatidae vs Hominidae ~20 Mya
Orangutans split Ponginae vs Homininae ~12–16 Mya
Gorillas split Gorilla vs Homo + Pan ~8–10 Mya
Chimps vs humans Pan vs Homo ~5–7 Mya

Principal Genera

Common Name Genera
Humans Homo
Chimps (common + bonobo) Pan
Gorillas Gorilla
Orangutans Pongo
Gibbons Hylobates
Baboons Papio
Lemurs (various) Lemur, Eulemur, etc.

Shared DNA

Humans are genetically closer to chimpanzees (Pan) than to gorillas (Gorilla). Table 3 shows approximate figures.These percentages refer to nucleotide identity in aligned genome regions. They vary slightly depending on how non-aligning or duplicated segments are treated.

Table 3: Shared DNA for selected species pairs.
Species Pair Shared DNA (%)
Human–Chimpanzee ~98.8%
Human–Gorilla ~98.4%
Human–Orangutan ~97%
Human–Gibbon ~96–97%
Human–Baboon ~93%
Human–Macaque ~92%
Human–Lemur ~80–85%
Chimpanzee–Gorilla ~98.3%
Chimpanzee–Orangutan ~97%
Gorilla–Orangutan ~96%
Chimpanzee–Gibbon ~95–96%

See also

Notes on other relevant VSI books.

VSI Details

  • Prehistory—A very short introduction
  • Chris Gosden
  • Volume 96
  • Published 2018

Appendix: Python Code to Make Map

import folium

# Create map centered roughly between UK and the Levant
m = folium.Map(location=[40, 25], zoom_start=4, tiles="OpenStreetMap")

# Define site data: (name, latitude, longitude, group)
sites = [
    # Near East
    ("Great Pyramid of Giza", 29.9792, 31.1342, "Egypt"),
    ("Aşıklı Höyük", 38.3439, 34.2631, "Central Anatolia"),
    ("Çayönü", 38.2806, 39.9714, "SE Turkey"),
    ("Göbekli Tepe", 37.2231, 38.9226, "SE Turkey"),
    ("Jericho (Tell es-Sultan)", 31.8703, 35.4433, "West Bank"),
    ("Ain Ghazal", 31.9511, 35.9457, "Jordan"),
    ("Beidha", 30.3292, 35.4658, "Jordan"),
    ("Abu Hureyra", 35.8925, 38.3850, "Syria"),
    ("Nevalı Çori", 37.5725, 38.6633, "SE Turkey"),
    
    # UK Sites
    ("Stonehenge", 51.1789, -1.8262, "UK"),
    ("Avebury", 51.4280, -1.8543, "UK"),
    ("Silbury Hill", 51.4154, -1.8545, "UK"),
    ("Grime’s Graves", 52.4842, 0.6493, "UK"),
    ("Ring of Brodgar", 59.0014, -3.2299, "Orkney"),
    ("Maeshowe", 58.9958, -3.1878, "Orkney"),
]

# Add markers to the map
for name, lat, lon, group in sites:
    folium.Marker(
        location=[lat, lon],
        popup=name,
        tooltip=name,
        icon=folium.Icon(color="blue" if group == "UK" else "green")
    ).add_to(m)

# Save map to HTML
m.save("prehistoric_sites_map.html")

References

1.
Daub, J.T., Moretti, S., Davydov, I.I., Excoffier, L., Robinson-Rechavi, M.: Detection of pathways affected by positive selection in primate lineages ancestral to humans. Molecular Biology and Evolution. 34, 1391–1402 (2017). https://doi.org/10.1093/molbev/msx083
2.
Bi, X., Zhou, L., Zhang, J.J., Feng, S., Hu, M., Cooper, D.N., Lin, J., Li, J., Wu, D.D., Zhang, G.: Lineage-specific accelerated sequences underlying primate evolution. Science Advances. 9, 1–12 (2023). https://doi.org/10.1126/sciadv.adc9507